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Another possible explanation is that mesangial cells express not only Nox4 but also Nox1, Nox2, and Nox5

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The p53 tumour suppressor protein, which is a transcription factor stabilized and activated in response to a range of cellular stresses including hyper-proliferation, DNA damage and shortage of glucose, was significantly up-regulated (by 5.9 fold) as compared to untreated cells (Supplementary Table S10)