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Later studies suggest that these are likely tonically active interneurons that release acetylcholine and glutamate and their firing is inhibited by both MORs and DORs (Ponterio et al., 2013

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doi: 10.1007/s00394-020-02188-2 [DOI] [PMC free article] [PubMed] [Google Scholar] 42.Nelson AJ, Peterson ED, Pagidipati NJ