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[15] The products of this reaction are pyruvate (the carbon skeleton of alanine) and glutamate: Alanine + α-Ketoglutarate ⇄ Glutamate + Pyruvate In a subsequent reaction catalyzed by glutamate dehydrogenase (EC 1.4.1.2), an enzyme located in the mitochondrial matrix, glutamate is deaminated to produce an ammonium ion (NH 4 + ), which enters the urea cycle, and α-ketoglutarate, which may re-enter the citric acid cycle

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